A paradigm can take hold even if incomplete, provided its core idea is sufficiently useful. For instance, Charles Darwin’s theory of natural selection offered a single principle that could explain the diversity of living species even though it still lacked an explanation for how traits actually passed from parent to offspring. In the late 1860s, he posited the missing mechanism, the erroneous notion of “pangenesis.” In it, he speculated that every cell in the body sheds tiny particles called “gemmules” that collect in the reproductive organs and transmit traits to offspring. Despite this error, Darwin’s core vision survived, and spread amongst biologists, before genetics supplied the necessary, physical mechanisms.
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